HSA Human Sheddome Atlas

Browse proteases

369 proteases with experimental cleavage evidence across the atlas.

Protease Gene / alias MEROPS Sources Substrates Sites Example substrates
merops+uniprotuniprot 1429 1429 ACE , ACE2 , ACP2 , ACP4 , ACVR1 , ACVR1B , ACVR1C , ACVR2A , ACVR2B , ACVRL1 , +1419
XS26.001 XS26.001 meropsmerops+uniprot 174 174 ADAM17 , ADAM32 , ADGRG1 , ADGRG3 , ALPP , APP , BAMBI , BCAM , BOC , BTN3A3 , +164
uniprot 91 91 ANO8 , ATAD3A , ATF6B , ATP5MF , ATP5MK , BAK1 , BCS1L , C3orf33 , CDAN1 , CERS1 , +81
M35.004 M35.004 merops 76 129 ABCA10 , ABCA6 , ABHD2 , ADAM29 , ADGRF1 , ADGRV1 , ALG9 , ASIC3 , ATP8A2 , ATRN , +66
CLE_M24 CLE_M24 merops 62 62 ABCD3 , ATAD3A , ATF6B , ATP5MF , ATP5MK , CDAN1 , CISD2 , CLIC1 , CLIC4 , CLPTM1 , +52
uniprot 58 58 ABCB10 , ABCB7 , ABCB8 , AFG3L2 , APOOL , ATP5MC1 , ATP5MC2 , ATP5MC3 , CCDC51 , CCDC90B , +48
hprd+uniprotpaper+uniprotuniprot 57 62 ACE , ADAM9 , ADGRB1 , ADGRF5 , ADGRG6 , ASAH2 , CD40LG , CLSTN1 , CORIN , DBH , +47
MBTPS1 autolysis P02.001, S08.063 hprd+uniprotmeropsmerops+uniprotuniprot 43 50 ADGRB2 , ADGRD1 , ADGRE2 , ADGRE3 , ADGRE4P , ADGRE5 , ADGRF1 , ADGRF3 , ADGRF5 , ADGRG1 , +33
PLG S01.151, S01.233 hprd+meropshprd+merops+pmapmeropsmerops+pmap 42 91 ABCC12 , APMAP , APP , ATP1B3 , BSG , CANX , CD81 , CLIC1 , CLIC4 , CNNM4 , +32
MEP1B M12.004 merops 36 52 ADAM10 , ADAM15 , ADAM9 , ANKAR , APLP2 , APP , ASPH , CANX , CD99 , CLIC1 , +26
FURIN furin S08.071 hprd+merops+pmaphprd+merops+pmap+uniprotmeropsmerops+uniprotuniprot 36 41 ADAM10 , ADAM12 , ADAM15 , ADAM19 , ADAM8 , ADGRB2 , ADGRF5 , ATP6AP1 , BACE1 , CNMD , +26
S01.280 S01.280 merops 35 57 ABCA3 , ADAM30 , AJAP1 , ALG6 , ATP2C1 , B3GALT1 , BLTP1 , CCDC47 , CDS2 , CHST4 , +25
PCSK5 S08.076 merops 28 29 ADAM10 , ADAM12 , ADAM17 , ADAM19 , ADAM8 , BACE1 , EDA , FURIN , HBEGF , INSR , +18
PCSK4 S08.074 merops 27 28 ADAM10 , ADAM12 , ADAM17 , ADAM19 , ADAM8 , BACE1 , EDA , FURIN , HBEGF , INSR , +17
PCSK6 S08.075 merops 27 28 ADAM10 , ADAM12 , ADAM17 , ADAM19 , ADAM8 , BACE1 , EDA , FURIN , HBEGF , INSR , +17
PCSK7 S08.077 merops 27 28 ADAM10 , ADAM12 , ADAM17 , ADAM19 , ADAM8 , BACE1 , EDA , FURIN , HBEGF , INSR , +17
MMP2 M10.003 hprd+merops+pmapmerops 26 32 ADGRG1 , APMAP , APP , CADM1 , CLIC1 , CLIC4 , CSPG4 , EPHB1 , FGFR1 , GRM4 , +16
PCSK2 S08.073 merops 24 25 ADAM10 , ADAM17 , ADAM19 , ADAM8 , BACE1 , EDA , FURIN , HBEGF , INSR , ITGA3 , +14
MEP1A M12.002 merops 21 30 ADAM9 , APP , ASPH , CLIC1 , CLSTN1 , DSC3 , DSG3 , DSG4 , F11R , FAT2 , +11
uniprot 19 20 AFG3L2 , ALPP , BST2 , BTC , CLN3 , EREG , NAGPA , PTGIR , RAET1G , SLC25A1 , +9
GZMB S01.010 hprd+meropsmerops 18 23 CACNA1C , CHRNA7 , CLIC1 , CLIC2 , CNGA1 , DLL1 , DSG2 , F2R , GRIA3 , GRIA4 , +8
ADAM17 M12.217 hprd+merops+pmapmeropsmerops+pmapuniprot 16 27 ACE , APP , AREG , BTC , CLSTN2 , CLSTN3 , ERBB4 , EREG , FCGR3A , GP1BA , +6
CLE_C14 CLE_C14 merops 13 13 CELSR2 , CYB5B , LRCH4 , MAN1A1 , MIA3 , NOTCH4 , OSBPL8 , SEMA4C , SLC25A46 , SLC4A3 , +3
KLK4 S01.251 merops 13 13 BACE1 , CDH1 , CDH2 , CDH3 , CDH4 , CDH5 , DSG1 , DSG3 , EPCAM , F2RL1 , +3
MMP14 M10.014 hprd+meropshprd+merops+pmapmeropsmerops+papermerops+pmapuniprot 12 32 APP , BCAM , BSG , CD44 , CDH1 , HBEGF , ITGAV , MMP14 , PTK7 , SDC1 , +2
MMP7 M10.008 meropsmerops+papermerops+pmap 12 18 ADAM28 , CDH1 , FAS , FASLG , IGHG1 , IGHG3 , IGHG4 , ITGB4 , MAG , OCLN , +2
S01.269 S01.269 merops 10 17 CANX , CD81 , CLIC1 , CSPG4 , IGHG1 , ITGAE , NNT , SELE , SLC25A29 , VDAC1
CLE_UNK CLE_UNK merops 8 9 ATP1A1 , COX4I1 , CYC1 , ITGA6 , LETM1 , NDUFB11 , ST14 , UQCRFS1
M12.033 M12.033 merops 8 8 CANX , CLIC4 , EBP , FKBP8 , MT-CO2 , NRXN3 , TMEM151A , VDAC3
ADAM10 M12.210 meropsmerops+pmapuniprot 8 13 APP , CD40LG , CLSTN2 , CLSTN3 , FASLG , FCER2 , TGFA , TNF
ELANE S01.131 hprd+meropshprd+merops+pmapmeropsuniprot 8 19 ATP1B3 , CDH5 , F2R , F2RL1 , GSDMB , IGHG1 , MICOS13 , SCNN1G
CTSG S01.133 hprd+meropshprd+merops+pmapmeropsmerops+pmapuniprot 8 17 CDH5 , F2R , F2RL1 , F2RL3 , GP1BA , IGHG1 , SELPLG , VDAC1
F2 S01.217 hprd+meropsmeropsmerops+pmap 7 9 CLIC1 , F2R , F2RL1 , F2RL2 , F2RL3 , GP5 , SDC4
CASP3 C14.003 merops 7 11 BAX , BCL2 , BCL2L1 , CD247 , LRBA , MCL1 , PLA2G6
furin-like protease uniprot 7 8 ADAM9 , ATP6AP2 , GLDN , LFNG , MMP23B , NOTCH1 , NOTCH3
CTSL C01.032 meropsuniprot 6 6 AADAC , ABCA3 , CLIC1 , GOLM2 , LRBA , PCDH15
METAP1D M24.028 meropsmerops+uniprot 6 6 DNAJC19 , MAOB , SFXN1 , SLC25A4 , SLC25A5 , VDAC1
KLK5 S01.017 meropsmerops+pmap 6 13 DSG1 , F2R , F2RL1 , F2RL3 , MEP1A , MEP1B
KLK6 S01.236 meropsmerops+pmap 5 16 APP , DSG1 , F2R , F2RL1 , F2RL3
CTSB C01.060 merops 5 12 APP , CLCA4 , MCOLN1 , PCNX3 , TMEM259
MMP9 M10.004 meropsmerops+uniprotuniprot 5 10 APP , DAG1 , NINJ1 , SDC1 , SDC4
M12.001 M12.001 merops 5 5 ARMH4 , CANX , NLGN3 , PCDHB5 , SELL
CASP1 C14.001 merops 5 5 BCL2L1 , GSDMD , MGAM2 , ST14 , VLDLR
thrombin uniprot 4 5 ADGRA2 , C9 , F2RL2 , SLC10A4
ADAM8 M12.208 meropsmerops+pmapuniprot 4 8 APP , CHL1 , FCER2 , TNF
MMP3 M10.005 meropsmerops+pmap 4 4 HBEGF , IGHG1 , IGHG3 , IGHG4
C01.073 C01.073 merops 4 10 CD209 , CLEC4M , FCER2 , OCLN
CAPN1 C02.001 hprd+meropshprd+merops+pmapmerops 4 13 BAX , BCL2L1 , F2R , F2RL1
KLK14 S01.029 meropsmerops+pmap 4 9 DSG1 , F2R , F2RL1 , F2RL3
uniprot 3 3 F2RL1 , F2RL2 , F2RL3
A02.001 A02.001 merops 3 4 APP , CLIC1 , CNNM4
PS1 uniprot 3 4 CREB3 , CREB3L3 , CREB3L4
uniprot 3 3 SDC2 , SDC3 , TREM2
MMP17 M10.017 merops 3 5 APP , HBEGF , TNF
ADAM19 M12.214 merops 3 4 ADAM19 , KITLG , TNF
LGMN C13.004 meropsuniprot 3 9 CD74 , IGHG1 , MPEG1
C01.033 C01.033 merops 3 3 IGHG2 , IGHG3 , IGHG4
CTSS C01.034 merops 3 15 CD74 , IGF2R , SCAI
PRSS3 S01.174 merops 3 3 APP , F2R , F2RL3
BACE1 A01.004 meropsmerops+paper 3 6 APP , BACE1 , SELPLG
CASP6 C14.005 hprd+meropsmerops 3 3 APP , LRBA , TEX2
PSEN1 A22.001 meropsuniprot 3 3 APP , CD44 , FLT1
GZMA S01.135 merops 3 3 CALHM1 , F2R , TMPO
PRTN3 S01.134 hprd+meropshprd+merops+pmapmerops 3 13 F2R , F2RL1 , PROCR
ST14 S01.302 hprd+meropsmerops 3 3 CDCP1 , F2RL1 , ST14
PARL S54.009 meropsmerops+uniprotuniprot 3 3 PARL , PGAM5 , PINK1
ADAM33 M12.244 merops 2 2 APP , TNFSF11
proprotein convertases uniprot 2 2 PLXNB1 , PLXNB2
HTRA2 S01.278 merops 2 5 APP , HTRA2
IDE M16.002 merops 2 21 APP , TGFA
ADAM9 M12.209 meropsmerops+pmap 2 2 APP , TNF
C25.002 C25.002 meropsmerops+pmap 2 5 C5AR1 , TNF
C66.001 C66.001 merops 2 2 IGHG1 , IGHG3
M64.001 M64.001 merops 2 2 IGHA1 , IGHA2
CTSD A01.009 hprd+meropsmerops 2 11 APP , DCD
BACE2 A01.041 hprd+merops+pmapmerops 2 11 APP , BACE2
M12.167 M12.167 merops 2 2 GP1BA , SELPLG
M10.039 M10.039 merops 2 3 FURIN , MMP14
caspase-3 uniprot 2 3 BCL2 , MCL1
S.pyogenes SpeB uniprot 2 3 CDH1 , GSDMA
a metalloproteinase uniprot 2 2 CDH1 , EPHB2
uniprot 2 2 B4GALT1 , CX3CL1
KLK1 S01.160 merops 2 3 DSG1 , F2R
TPSB2 S01.143 hprd+meropsmerops+pmap 2 5 F2R , F2RL1
S01.001 S01.001 merops 2 3 CADM1 , F2R
CLE_S26 CLE_S26 merops 2 2 CANX , SSR2
XT01.001 XT01.001 merops 2 20 DCT , PMEL
trypsin uniprot 2 2 F2RL1 , MPEG1
CAPN2 C02.002 hprd+meropsmerops 2 9 BAX , F2RL1
C11.001 C11.001 merops 2 4 BACE1 , IL20RA
uniprot 2 2 ERVPABLB-1 , ERVS71-1
RHBDL4/RHBDD1 uniprot 2 2 BIK , STEAP3
uniprot 1 1 GGT3P
uniprot 1 1 SLN
uniprot 1 1 NRG2
uniprot 1 1 ADGRB1
furin and PCSK7 uniprot 1 1 ADAM10
uniprot 1 1 EREG
uniprot 1 1 TMPRSS2
uniprot 1 1 TMPRSS11D
M41.003 M41.003 merops 1 2 OPA1
uniprot 1 1 OPA1
uniprot 1 2 OPA1
PRCP S28.001 merops 1 1 GPR37L1
KLK13 S01.306 merops 1 1 CDH16
SPPL3 uniprot 1 1 CLN5
uniprot 1 1 CLN5
uniprot 1 1 ATP6AP2
uniprot 1 1 ATP6AP2
uniprot 1 1 CNMD
ADGRL1 P02.010 merops 1 1 ADGRL1
uniprot 1 1 CLSTN1
uniprot 1 2 EGF
CPM M14.006 merops 1 1 EGF
uniprot 1 1 TGFA
SPPL2B A22.004 merops 1 2 TNF
SPPL2A A22.007 merops 1 2 TNF
SPPL2A or SPPL2B uniprot 1 2 TNF
uniprot 1 1 TNF
uniprot 1 1 TNF
uniprot 1 1 TNF
C25.001 C25.001 merops+pmap 1 2 TNF
C25.003 C25.003 merops+pmap 1 1 TNF
uniprot 1 1 PIGR
C01.001 C01.001 merops 1 2 IGHG1
MMP12 M10.009 merops 1 1 IGHG1
PGA4;PGA5 A01.070 merops 1 1 IGHG1
PGA3;PGA4 A01.001 merops 1 1 IGHG1
M26.001 M26.001 merops 1 1 IGHA1
S06.007 S06.007 merops 1 1 IGHA1
U9G.049 U9G.049 merops 1 1 IGHA2
M9G.002 M9G.002 merops 1 1 GYPA
uniprot 1 1 C9
trypsin; to produce soluble form uniprot 1 1 TFRC
uniprot 1 1 TFRC
uniprot 1 2 CD74
A32.002 A32.002 merops 1 1 RPN1
caspases uniprot 1 2 APP
uniprot 1 1 APP
C47.003 C47.003 merops 1 2 APP
MMP16 M10.016 merops 1 4 APP
S9G.058 S9G.058 merops 1 2 APP
uniprot 1 1 APP
beta-secretase uniprot 1 1 APP
HTRA1 S01.277 merops 1 3 APP
U9G.071 U9G.071 merops 1 3 APP
caspase-6; when associated with variant 670-N-L-671 uniprot 1 1 APP
MME M13.001 merops 1 2 APP
ECE1 M13.002 merops 1 4 APP
S01.243 S01.243 merops 1 4 APP
ACE M02.001 meropsuniprot 1 3 APP
U9G.055 U9G.055 merops 1 1 APP
S01.422 S01.422 merops 1 5 APP
APEH S09.004 merops 1 3 APP
BLMH C01.084 merops 1 3 APP
M9G.026 M9G.026 merops 1 3 APP
C04.001 C04.001 merops 1 1 APP
PITRM1 M16.009 merops 1 9 APP
A01.035 A01.035 merops 1 1 APP
MMEL1 M13.008 merops 1 2 APP
uniprot 1 1 APP
alpha-secretase uniprot 1 1 APP
CTSE A01.010 merops 1 2 APP
theta-secretase uniprot 1 1 APP
uniprot 1 1 APP
A22.015 A22.015 merops 1 2 APP
uniprot 1 1 APP
gamma-secretase; site 1 uniprot 1 1 APP
PIGK C13.005 merops 1 1 ALPP
uniprot 1 1 ALPP
HPN S01.224 merops+pmap 1 1 HPN
uniprot 1 1 HPN
uniprot 1 1 INSR
uniprot 1 1 INSR
uniprot 1 1 ITGAV
uniprot 1 1 ITGAV
uniprot 1 1 GP1BA
uniprot 1 1 IGF1R
uniprot 1 1 IGF1R
uniprot 1 1 ITGA2B
uniprot 1 1 ITGA2B
uniprot 1 1 ITGA2B
uniprot 1 1 ITGA5
uniprot 1 1 IL6R
ADAM10 and ADAM17 uniprot 1 1 IL6R
uniprot 1 1 CSF1
uniprot 1 1 CSF1
uniprot 1 1 LCT
uniprot 1 1 FURIN
uniprot 1 1 GHR
uniprot 1 1 CDH1
CTSA S10.002 merops 1 1 LAMP2
uniprot 1 1 SI
uniprot 1 1 INSRR
uniprot 1 1 INSRR
uniprot 1 1 AREG
uniprot 1 1 CD1E
MUC1 S71.001 merops 1 1 MUC1
uniprot 1 1 MUC1
uniprot 1 1 ABO
HM13/SPP uniprot 1 1 XBP1
uniprot 1 1 XBP1
uniprot 1 2 TNFRSF1A
uniprot 1 1 GGT1
uniprot 1 1 TNFRSF1B
CMA1 S01.140 merops 1 1 KITLG
uniprot 1 1 KITLG
CTRB1;CTRB2 S01.152 merops 1 4 C5AR1
uniprot 1 1 ZP3
PLAU S01.231 merops 1 1 ITGA6
PLAU in invasive prostate cancer uniprot 1 1 ITGA6
uniprot 1 1 ITGA6
uniprot 1 1 ITGA6
uniprot 1 1 ITGA6
Staphylococcus aureus/SspP uniprot 1 1 CXCR2
MMP1 M10.001 merops 1 1 F2R
F10 S01.216 merops 1 1 F2R
PROC S01.218 merops 1 1 F2R
thrombin and CTSG uniprot 1 1 F2R
uniprot 1 1 ITGA3
uniprot 1 1 ITGA3
uniprot 1 1 DPP4
uniprot 1 1 EPHB2
M08.001 M08.001 merops 1 1 ABCC1
uniprot 1 1 BTC
uniprot 1 1 GGT5
uniprot 1 2 SCNN1A
uniprot 1 1 ITGAE
uniprot 1 1 ITGAE
XT01.003 XT01.003 merops 1 2 DCT
ERAP2 M01.024 merops 1 4 PMEL
uniprot 1 1 PMEL
furin-like proprotein convertase uniprot 1 1 PMEL
uniprot 1 1 PMEL
ADAM metalloproteinases uniprot 1 1 PMEL
uniprot 1 2 TMPO
uniprot 1 1 TMPO
uniprot 1 1 NOTCH1
uniprot 1 1 UQCRFS1
uniprot 1 1 FASLG
uniprot 1 1 ADGRE5
CASP2 C14.006 merops 1 1 LRBA
PRSS8 uniprot 1 1 SCNN1G
CELA1 S01.153 merops 1 1 SCNN1G
uniprot 1 1 ITGA8
uniprot 1 1 CACNA2D1
CTSK C01.036 merops 1 1 ATP1B3
C01.072 C01.072 merops 1 1 F2RL1
PRSS1 S01.127 merops+pmap 1 1 F2RL1
M04.005 M04.005 merops 1 2 F2RL1
uniprot 1 1 F2RL1
uniprot 1 1 F2RL1
uniprot 1 1 NRG3
CASP3 or CASP7 uniprot 1 1 GSDMD
uniprot 1 1 GSDMD
enterovirus 71 (EV71) Protease 3C and coronavirus SARS-CoV-2 proteinase nsp5 uniprot 1 1 GSDMD
caspases CASP1, CASP4, CASP5 and CASP8 uniprot 1 1 GSDMD
uniprot 1 1 GSDMD
papain uniprot 1 1 GSDMD
uniprot 1 1 GSDMD
uniprot 1 1 ZP1
uniprot 1 1 DLK1
GZMM S01.139 merops 1 1 DCD
uniprot 1 1 DCD
uniprot 1 1 DCD
uniprot 1 1 TMPRSS15
PKD1L2 P02.038 merops 1 1 PKD1
MMP8 M10.002 merops 1 1 EFNB1
uniprot 1 1 EFNB1
uniprot 1 1 DHODH
uniprot 1 1 NRG1
uniprot 1 1 NOTCH2
uniprot 1 1 MST1R
uniprot 1 1 MST1R
PLAT S01.232 merops 1 1 GRIN1
ASTL uniprot 1 1 ZP2
caspase-1 uniprot 1 1 BCL2L1
uniprot 1 1 LRP1
uniprot 1 1 MGAT5
uniprot 1 1 GALNT2
uniprot 1 1 GALNT1
uniprot 1 1 BST2
uniprot 1 1 ZP4
urokinase uniprot 1 1 ITGA7
uniprot 1 1 ITGA7
uniprot 1 1 ITGA7
uniprot 1 1 ITGA7
DAG1 S72.001 merops 1 1 DAG1
uniprot 1 1 DAG1
DDI2 uniprot 1 1 NFE2L1
uniprot 1 1 NFE2L1
CLCA4 M87.002 merops 1 1 CLCA4
S08.025 S08.025 merops 1 2 OCLN
uniprot 1 1 PRSS8
uniprot 1 1 PRSS8
uniprot 1 1 PRSS8
uniprot 1 2 MPEG1
uniprot 1 1 GNPTAB
ASPRV1 A28.004 meropsmerops+uniprot 1 3 ASPRV1
uniprot 1 1 ADGRF1
uniprot 1 1 RAET1G
uniprot 1 1 ADGRD1
uniprot 1 2 GLDN
BMP1 uniprot 1 1 GLDN
uniprot 1 1 CACNA2D4
furin like-convertase uniprot 1 1 ADGRG6
uniprot 1 4 CD163
uniprot 1 1 ADGRG3
lysosomal cysteine proteases uniprot 1 1 PLD3
uniprot 1 2 VSTM4
uniprot 1 1 SPACA3
uniprot 1 1 ADGRG5
uniprot 1 1 ADGRG4
uniprot 1 1 ADGRG2
uniprot 1 2 SMIM20
uniprot 1 1 SMIM20
CAPS3, CAPS6 and CAPS9 uniprot 1 1 GSDMB
granzyme A uniprot 1 2 GSDMB
uniprot 1 1 GSDMB
uniprot 1 1 NRG4
uniprot 1 1 STAB2
uniprot 1 1 ADGRV1
uniprot 1 1 PTPRN2
uniprot 1 1 NINJ1
S01.136 S01.136 merops 1 1 ABCC12
uniprot 1 1 SORCS2
uniprot 1 1 GSDMA
thrombin or trypsin uniprot 1 1 F2RL3
uniprot 1 1 HBEGF
uniprot 1 1 HBEGF
uniprot 1 1 HBEGF
uniprot 1 1 NOTCH4
M16.003 M16.003 merops 1 1 SDHC
uniprot 1 1 ABCA3
uniprot 1 2 ATF6B
uniprot 1 1 COL25A1
autolysis; required for cell surface localization and secretion uniprot 1 1 TMPRSS13
uniprot 1 1 TMPRSS13
uniprot 1 1 ACE2
ST14/MT-SP1 uniprot 1 1 CDCP1
uniprot 1 1 ERVMER34-1
ADGRL3 P02.011 merops 1 1 ADGRL3
paper+uniprot 1 1 CLTRN
M12.161 M12.161 merops 1 2 GP6
uniprot 1 1 NRXN3
uniprot 1 2 ITM2C
uniprot 1 1 TPSG1
uniprot 1 1 TPSG1
uniprot 1 1 ATAD3A
uniprot 1 1 CACNA2D2
CASP7 C14.004 merops 1 1 PTGFRN
uniprot 1 1 PISD
uniprot 1 1 LNPEP
uniprot 1 1 GGT7
uniprot 1 1 TMPRSS11E
uniprot 1 1 MGAT4A
uniprot 1 1 NOTCH3
uniprot 1 1 COL17A1
uniprot 1 1 COL17A1
uniprot 1 1 CLCA2
CLCA2 M87.003 merops 1 1 CLCA2
uniprot 1 1 ERVW-1
uniprot 1 1 ITM2B
uniprot 1 1 ITM2B
CORIN S01.019 merops 1 2 CORIN
uniprot 1 1 CORIN
uniprot 1 1 CORIN
uniprot 1 1 MMP24
uniprot 1 1 ADGRG1